The question below comprises four scattered segments of a paragraph. Identify from among the four choices the sequences that correctly assemble the segments and complete the paragraph.

1. It was long thought to be a distinct species, but genetic studies have shown it to be the southernmost subspecies of plains zebra.
2. It is considered particularly close to Burchell’s zebra.
3. And just as the Burchell’s, its name is derived from its call.
4. The quagga is an extinct subspecies of plains zebra that lived in South Africa until the 19th century.