Comprehension Passage

Direction: Study the following information and answer the following questions.

I * J means I is the father of J.

I ^ J means I is the mother of J.

I % J means I is the son of J.

I # J means I is daughter of J.

I ! J means I is the husband of J.

I ~ J means I is the wife of J.

Which of the following conclusions can be drawn from the expression?

Y * Z % V # U ! T # S ~ R

S is the grandmother of Y.

T is the mother-in-law of R.

Z is the grandson of S.

V is the granddaughter of R.

Z is the grandfather of U.