If 'A & B' means 'A is the brother of B', 'A # B' means 'A is the daughter of B', and 'A @ B' means 'A is the wife of B', then how is K related to Z in the following expression?

'Z @ R & T # H @ K'

1
Paternal uncle
2
Father-in-law
3
Paternal grandfather
4
Father

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