Study the following information carefully and answer the given question.

'A % B' means 'A is the mother of B', 'A # B' means 'A is the brother of B', 'A @ B' means 'A is the father of B' and 'A + B' means 'A is the husband of B'.

If (T @ P + Q % R # S), then how is T related to S?

1
Grandfather
2
Father
3
Grandmother
4
Brother

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