If 'A @ B' means 'A is the mother of B' and 'A % B' means 'A is the brother of B', then which of the following statements is correct for the given expression?

G @ P % U % V @ R

1
U is the father of R.
2
G is the mother of V.
3
R is the daughter of G.
4
U is the sister of R.

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