If A & B means A is the husband of B, A # B means B is the son of A, A @ B means B is the sister of A and A % B means B is the brother of A. brother, then which of the following expressions definitely means that S is the brother of T?   

1
P & Q # R @ S % T
2
Q & S # T @ R % P
3
R @ S % T # Q & P
4
Q & P # T % S @ R 

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