The following statements were made about the structure of the 30-nm chromatin fiber:

A. In the solenoid model, the linker DNA connects the consecutive core particles.

B. In the zig-zag model, alternating nucleosomes become interacting neighbors.

C. In the solenoid model, 12 nucleosomes are organized into two separate stacks, whereas 8 nucleosomes per turn make a single stack in the zig-zag model.

D. H1 histone is essentially required as per the zig-zag model, but not as per the solenoid model.

E. Chromatin fibers prepared with H4 histones that lack their tails could fold into higher-order fibers.

Which one of the following options represents the combination of all correct statements?