In the following question, the symbols ' + - ^ % are used with the following meaning as illustrated below.

A + B means 'A is husband of B'.

A - B means 'A is mother of B'.

A ^ B means 'A is the sister of B'.

A % B means 'A is brother of B'.

If L + M - N ^ R % Q, then how is L related to N?

1
Sister
2
Brother
3
Uncle
4
Father
5
Mother

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