The complexity of a food web in a community is quantified using certain parameters which are defined below. Which of the following is an INCORRECT representation?

1
Chain length = Average number of links between trophic levels / Total number of trophic levels
2
Connectance = actual number of links in a food web / potential number of links in a food web
3
Potential links in a food web where 'n' species are present = n(n-1) / actual number of links
4
Linkage density = actual number of links in a food web / Number of species in a food web

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