Which of the following correctly describes the malate-aspartate shuttle?

1
It generates FADH2 during the transport of reducing equivalents across the mitochondrial membrane.
2
It operates primarily in skeletal muscle and brain tissue.
3
It transports reducing equivalents from the cytosol to the mitochondrial matrix by converting oxaloacetate to malate.
4
 It generates NADH when oxaloacetate is converted back to malate in the cytosol.

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