In the sentence, certain words are in bold and numbered from A to E, which are the possible pairs to be interchanged. Choose the pair(s) of words which need to be interchanged to make the sentence grammatically correct and meaningful.

The last known (A)resting place of Homo erectus, one of the most (B)traced human ancestors and the first to walk fully (C)upright, has been (D) successful to a (E)floodplain near the longest river on the Indonesian island of Java.