What causes the formation of growth rings or annuli in cattle horns?

  1. Seasonal variations in temperature
  2. Changes in diet composition
  3. Periodic shedding of horn layers
  4. Annual reproductive cycles

Choose the most appropriate answer from the options given below: 

1
1 Only
2
2 Only 
3
3 Only
4
All of the above 
5
Question Not Attempted

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