Study the following information carefully and answer the given questions:

If ‘P # Q’ means ‘P is the mother of Q’

If ‘P % Q’ means ‘P is the father of Q’

If ‘P & Q’ means ‘P is the sister of Q’

If ‘P @ Q’ means ‘P is the brother of Q’

If ‘D @ E # C % O @ R’ is given, then how is E related to O?

1
Grandfather
2
Grandmother
3
Grandson
4
Granddaughter

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