Use the relations defined below solve the question.

A = B means 'A is the father of B'

A > B means 'A is the sister of B'

A < B means 'A is the wife of B'

A % B means 'A is the mother of B'

A $ B means 'A is the son of B'

What should come in place of the question mark (?) to establish, that A is the brother of D? A$B%C? D%E

1
>
2
$
3
<
4
=

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