Comprehension Passage

In these types of questions the symbols $, %, &, @ are used with the following meanings as illustrated below:

P $ Q means P is the sister of Q.

P % Q means P is the brother of Q.

P & Q means P is the mother of Q.

P @ Q means P is the son of Q.

How is A related to E in the given expression?

A % B $ C @ D & E

1
Brother.
2
Son.
3
Uncle.
4
None.

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