Expression ‘A $ B ! C @ D % E’ indicates some relations between A, B, C, D and E, where “A $ B means, A is a niece of B”, “A @ B means, A is a father of B”, “A % B means A is a son of B”, “A ! B means A is a sister of B”, "A # B means A is a mother of B". What does “#” mean in E # A?

1
Daughter
2
Mother
3
Daughter in law
4
Mother in law

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