If,

'C @ S' means 'C is the father of S',

'C * S' means 'C is the brother of S',

'C # S' means 'C is the sister of S',

'C $ S' means 'C is the daughter of S',

'C = S' means 'C is the mother of S',

then how is Z related to Q in the following expression?

Q $ R @ S * T $ Z