Which of the following letter-clusters should replace # and % so that the pattern and relationship followed between the letter-cluster pair on the left side of :: is the same as that on the right side of :: ?

# : USN :: PNI : %

1
# = LKP, % = MKJ
2
# = WUP, % = NLG 
3
# = MKI, % = NMJ 
4
# = MUP, % = NKJ

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