‘A + B’ means ‘B is the mother of A’

‘A # B’ means ‘B is the sister of A’

‘A @ B’ means ‘B is the brother of A’

‘A % B’ means ‘B is the daughter of A’

If A # B @ C % D + E, then how is E related to A? 

1
Wife 
2
Sister
3
Brother's wife
4
Brother's daughter

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