Comprehension Passage

Read the following instructions carefully and answer the following question. 

If 'A + B' means 'A is father of B'.

If 'A × B' means 'A' is sister of B'.

If 'A ÷ B' means 'A is wife of B'.

If 'A % B' means 'A is mother of B'.

If 'A * B' means 'A is son of B'.

What should come in place of question mark to establish 'Q is grandmother of T'?

P × Q ? R + S + T

1
%
2
+
3
÷
4
× 

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