If A @ B' means 'A is brother of B', 'A # B' means 'A is father of B', 'A $ B' means 'A is sister of B', 'A + B' means 'A is the mother of B' and 'A % B' means 'A is the husband of B', then which of the following expression represents P as the father of S?
1
P % Q $ R + S
2
P % Q + R @ S
3
P % Q + R # S
4
P % Q $ R $ S