What would be the effect of retinoic acid (RA) treatment on the ‘positional information’ of blastema cells present on the amputated newt limb?

1
RA will have no effect. 
2
The cells will become respecified to more proximal position. 
3
The cells will become respecified to more distal position. 
4
Cells will lose their positional information and remain as dedifferentiated cells 

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