Who explained - "Succession is an extraordinarily mobile phenomenon whose processes are not to be stated as fixed laws, but only a general principles of exceedingly broad nature and whose results need not and frequently do not occur in any definite predictable way"? 

1
F.E. Clement (Relay Floristics view)
2
F.E. Egler (Initial Floristics Composition view)
3
H.A. Gleason (Individualistic Concept)
4
D. Tilman (Resource-ratio hypothesis)

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